Understanding the origin and maintenance of biodiversity is a central challenge in ecology.Literature search found that global temperature changed cyclically but asymmetrically during Quaternary Glaciations.Slow global temperature decrease but rapid increase in every climate oscillation cycle is presented as an asymmetric temperature pattern:slow cooling but rapid warming.Asymmetric temperature changes have drived geographic distribution of communities to a pattern ofslow retreat and rapid expansion.Species in a community were isolated in the long term and were reassembled provisionally.As in many phylogeographic cases,genetic variations in general occurred inslow retreatstages.In rapid expansion stages,variations expanded in different routes and were recomposed in species populations.Our study suggests that a large-scale research platform for natural community reflectingslow retreat and rapid expansionshould be established.Combined with phylogeographic methods,origin and maintenance of community should be reconstructed in multi-scale and multi-dimension to predict the future of natural communities in climate change and human influence.